$QGEN

Wellington Management

12,406,763 股

Steve Cohen - Point72 Asset Management

1,588,845 股

Israel Englander - Millennium Management

3,845,605 股

Fidelity International (FIL)

1,290,558 股

Amundi

3,683,362 股

UBS Group

3,419,195 股

Ken Griffin - Citadel Advisors

832,391 股

Dmitry Balyasny - Balyasny Asset Management

176,542 股

Morgan Stanley

5,042,391 股

Nuveen

1,161,094 股

Voya Investment Mgmt

186,616 股

Adage Capital Partners

310,000 股

Cliff Asness - AQR Capital Management

426,739 股

PSP Investments

118,503 股

Northern Trust Corp

1,018,165 股

Larry Fink - BlackRock

5,905,678 股

Jan van Eck - VanEck

114,087 股

SG Americas Securities

113,908 股

State Street Corp

2,183,206 股

Geode Capital Management

1,074,640 股

Lazard Asset Management

36,510 股

Jamie Dimon - JPMorgan Chase

895,404 股

David E. Shaw - D.E. Shaw & Co.

74,291 股

Invesco

584,091 股

Franklin Resources

206,443 股

Jane Street Group

63,694 股

Legal & General

108,401 股

Neuberger Berman

28,715 股

Charles Schwab

99,427 股

Two Sigma Investments

14,317 股

PNC Financial Services

10,659 股

Mitsubishi Ufj Asset Management Co., Ltd.

6,540 股

Capital International Investors

0 股

Jefferies Group

618,798 股

Exoduspoint Capital Management, Lp

321,161 股

Verition Fund Management LLC

163,845 股

Louis Moore Bacon - Moore Capital Management

0 股

New York State Teachers Retirement System - New York State Teachers Retirement System

0 股

Nomura Holdings INC

0 股

John Hussman - Hussman Strategic Advisors

0 股