$TSHA

David Solomon - Goldman Sachs

22,369,645 股

Israel Englander - Millennium Management

3,281,820 股

David E. Shaw - D.E. Shaw & Co.

4,427,301 股

SG Americas Securities

2,176,742 股

Morgan Stanley

17,965,371 股

Steve Cohen - Point72 Asset Management

461,856 股

Renaissance Technologies

413,203 股

Two Sigma Investments

742,081 股

Voya Investment Mgmt

511,983 股

Ensign Peak Advisors

275,111 股

State Street Corp

11,209,078 股

Geode Capital Management

5,531,955 股

Larry Fink - BlackRock

18,365,608 股

Ken Griffin - Citadel Advisors

411,135 股

Charles Schwab

1,808,599 股

Northern Trust Corp

1,814,318 股

Dmitry Balyasny - Balyasny Asset Management

83,019 股

Nuveen

549,153 股

Wellington Management

626,361 股

UBS Group

517,698 股

BNP Paribas

120,200 股

Cliff Asness - AQR Capital Management

177,309 股

AllianceBernstein (AB)

189,240 股

Invesco

368,021 股

Mitsubishi Ufj Asset Management Co., Ltd.

56,577 股

Janus Henderson Group

69,000 股

Franklin Resources

108,399 股

Legal & General

104,308 股

T. Rowe Price

158,954 股

Neuberger Berman

10,823 股

PNC Financial Services

1,043 股

Canada Pension Plan Investment Board

0 股

Jane Street Group

0 股

Adage Capital Partners

0 股

Jefferies Group

479,000 股

Polen Capital Management

319,748 股

Paul Tudor Jones - Tudor Investment Corp

143,575 股

Nomura Holdings INC

0 股

Louis Moore Bacon - Moore Capital Management

328,831 股

Jamie Dimon - JPMorgan Chase

0 股