$QGEN

Wellington Management

14,364,303 股

Fidelity International (FIL)

1,577,141 股

Steve Cohen - Point72 Asset Management

511,912 股

Dmitry Balyasny - Balyasny Asset Management

433,592 股

Ken Griffin - Citadel Advisors

531,381 股

Morgan Stanley

4,190,913 股

Israel Englander - Millennium Management

597,837 股

Voya Investment Mgmt

195,596 股

Capital International Investors

2,436,169 股

Adage Capital Partners

285,000 股

Northern Trust Corp

1,006,157 股

Cliff Asness - AQR Capital Management

353,579 股

Janus Henderson Group

366,661 股

Lazard Asset Management

80,718 股

Jane Street Group

160,639 股

Invesco

725,508 股

Larry Fink - BlackRock

5,250,461 股

State Street Corp

2,089,631 股

Geode Capital Management

1,032,837 股

AllianceBernstein (AB)

138,459 股

Franklin Resources

170,703 股

Two Sigma Investments

42,438 股

Neuberger Berman

29,956 股

David E. Shaw - D.E. Shaw & Co.

19,397 股

Legal & General

69,526 股

Charles Schwab

91,804 股

UBS Group

0 股

Nuveen

0 股

PNC Financial Services

0 股

Jan van Eck - VanEck

0 股

Ensign Peak Advisors

0 股

Verition Fund Management LLC

65,075 股

Nomura Holdings INC

44,839 股

PSP Investments

110,980 股

New York State Teachers Retirement System

12,711 股

Segall Bryant & Hamill, LLC

0 股

Louis Moore Bacon - Moore Capital Management

20,000 股

Jamie Dimon - JPMorgan Chase

0 股